Satu Ramula profile picture
Docent, Ecology and Evolutionary Biology
University Lecturer, Ecology and Evolutionary Biology
PhD, adjunct professor


+358 29 450 4227
+358 50 465 3149
Vesilinnantie 5

Areas of expertise

Demographic methods
invasive species
plant ecology
population ecology
structured population models


2020-  University lecturer in plant ecology, University of Turku

2015-2020 Academy of Finland research fellow, University of Turku

2006 PhD, Stockholm University


  • Current courses:
  • BIOL1007 Structure and function of plants
  • BIOL4220 Biological data analysis: multivariate methods
  • EKOL2108 Population ecology
  • EKOL2103 Laboratory practicals in ecology
  • EKOL2354 Identification of vascular plants in Finland
  • EKOL2302 Terrestial plant ecology, field course
  • MAAN6154 Biogeography
  • EKOL4101 Ecological interactions


My research focuses on population ecology and invasive species. I am broadly interested in population-level ecological, evolutionary and methodological questions, including drivers of plant invasions, species interactions, and data quality issues. I often apply structured population models to understand spatial and temporal variation in wild populations, and to produce management recommendations. I currently lead the project on 'Going belowground to curb plant invasions' funded by the Academy of Finland.

I am a member of the science committee for the COMPADRE & COMADRE database ( project to create an open-access database of matrix population models for both animals and plants from all over the world to facilitate its usage for scientific and teaching purposes. These databases are a repository of demographic data from (mostly) published literature. As the number of published matrix population models (MPMs) grows, so do the opportunities to ask exciting questions about plant and animal demography, ecology, and evolution at a global and cross-taxonomic scale. The underlying motivation behind COMPADRE and COMADRE is to facilitate this research by collecting, digitising, and archiving MPMs at a central repository in a standardised format.  




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Phenotypic plasticity masks range-wide genetic differentiation for vegetative but not reproductive traits in a short-lived plant (2021)

Ecology Letters
Villellas Jesus, Ehrlén Johan, Crone Elizabeth E., Csergo Anna Mária, Garcia Maria B., Laine Anna-Liisa, Roach Deborah A., Salguero-Gómez Roberto, Wardle Glenda M., Childs Dylan Z., Elderd Bret D., Finn Alain, Munné-Bosch Sergi, Bachelot Benedicte, Bódis Judit, Bucharova Anna, Caruso Christina M., Catford Jane A., Coghill Matthew, Compagnoni Aldo, Duncan Richard P., Dwyer John M., Ferguson Aryana, Fraser Lauchlan H., Griffoul Emily, Groenteman Ronny, Hamre Liv Norunn, Helm Aveliina, Kelly Ruth, Laanisto Lauri, Lonati Michele, Münzbergová Zuzana, Nuche Paloma, Olsen Siri Lie, Oprea Adrian, Pärtel Meelis, Petry William K., Ramula Satu, Rasmussen Pil U., Enri Simone Ravetto, Roeder Anna, Roscher Christiane, Schultz Cheryl, Skarpaas Olav, Smith Annabel L., Tack Ayco J.M., Töpper Joachim Paul, Vesk Peter A., Vose Gregory E., Wandrag Elizabeth, Wingler Astrid, Buckley Yvonne M.
(Vertaisarvioitu alkuperäisartikkeli tai data-artikkeli tieteellisessä aikakauslehdessä (A1))

Global gene flow releases invasive plants from environmental constraints on genetic diversity (2020)

Proceedings of the National Academy of Sciences of the United States of America
Annabel L. Smith, Trevor R. Hodkinson, Jesus Villellas, Jane A. Catford, Anna Mária Csergo, Simone P. Blomberg, Elizabeth E. Crone, Johan Ehrlén, Maria B. Garcia, Anna-Liisa Laine, Deborah A. Roach, Roberto Salguero-Gómez, Glenda M. Wardle, Dylan Z. Childs, Bret D. Elderd, Alain Finn, Sergi Munné-Bosch, Maude E. A. Baudraz, Judit Bódis, Francis Q. Brearley, Anna Bucharova, Christina M. Caruso, Richard P. Duncan, John M. Dwyer, Ben Gooden, Ronny Groenteman, Liv Norunn Hamre, Aveliina Helm, Ruth Kelly, Lauri Laanisto, Michele Lonati, Joslin L. Moore, Melanie Morales, Siri Lie Olsen, Meelis Pärtel, William K. Petry, Satu Ramula, Pil U. Rasmussen, Simone Ravetto Enri, Anna Roeder, Christiane Roscher, Marjo Saastamoinen, Ayco J. M. Tack, Joachim Paul Töpper, Gregory E. Vose, Elizabeth M. Wandrag, Astrid Wingler, Yvonne M. Buckley
(Vertaisarvioitu alkuperäisartikkeli tai data-artikkeli tieteellisessä aikakauslehdessä (A1))

Comments to "Persistent problems in the construction of matrix population models" (2020)

Ecological Modelling
Che-Castaldo Judy, Jones Owen R., Kendall Bruce E., Burns Jean H., Childs Dylan Z., Ezard Thomas H.G., Hernandez-Yanez Haydee, Hodgson David J., Jongejans Eelke, Knight Tiffany, Merow Cory, Ramula Satu, Stott Iain, Vindenes Yngvild, Yokomizo Hiroyuki, Salguero-Gómez Roberto
(Kirjoitus tai data-artikkeli tieteellisessä aikakauslehdessä (B1))